Local PhD programs
- Leiden University and Leiden University Medical Center, The Netherlands
  - PhD courses Leiden University
  - PhD courses Leiden University Medical Center (Boerhaave)
- Medical Genetics Centre (MGC)

- Johannes Gutenberg University, Germany
- PhD program

- Imperial College London, United Kingdom
  - Postgradute courses

- Helmholtz-Zentrum für Infektionsforschung, Germany
- PhD program

- University of Cologne, Germany
  - Postgraduate studies

- University College London, United Kingdom
  - PhD program

- University of Oxford, United Kingdom
  - Postgraduate program

- Institut für Molekulare Biotechnologie
- PhD program

Other training facilities
- United Kingdom: Wellcome Trust Advanced Courses Programme

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